Marcelo Pires

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(949) 582-4809
Biological Sciences Instructor
Math, Science and Engineering
Saddleback College


2007 - Ph.D. Ecology and Evolutionary Biology
University of California, Riverside
Advisor: Dr. David N. Reznick
Dissertation Title: The Evolution of Placentas in Poeciliid Fishes

2001 - M.Sc., Biology
California State Polytechnic University, Pomona
Advisor: Dr. Glenn R. Stewart
Thesis Title: Allocation of Reproductive Output in the Western Pond Turtle (Clemmys marmorata) in Southern California

1998 - B.Sc., Biology. Summa cum laude
California State Polytechnic University, Pomona

1994-1995 – Early undergraduate work, Biology
Federal University of Rio de Janeiro, Brazil (UFRJ)


Depending on the semester, I may teach Principles of Cellular Biology (General Biology I - Bio 4AH), Principles of Organismal Biology (General Biology II, Bio 4BH), General Biology for non-bio majors (Bio 20), Animal Behavior (Bio 43), Human Evolution (Bio 40), Human Biology (Bio 30), Human Anatomy (Bio 11/110), Human Physiology (Bio 12/120), or Biology of Human Disease (Bio 45). I also teach a study abroad program in Brazil or Costa Rica (Bio 230, Bio 231, or Bio 234) during summer.


Nunes, C.E.P., Yasseen, K., Luxon, R., Williams, H., Koehler, S., Pires, M. 2018. Orchids of the State Park of Serra do Mar, São Paulo, Brazil Núcleo Santa Virgínia. The Field Museum of Natural History Field Guides. Field Guide #1000.

Pires, M. N. and D. N. Reznick. 2018. Life-history evolution in the fish genus Poecilia (Poeciliidae: Cyprinodontiformes: subgenus Pamphorichthys): an evolutionary origin of extensive matrotrophy decoupled from superfetation. Biological Journal of the Linnean Society 125(3): 547–560.  Click here for the article.

Pires, M. N., A. I. Banet, B. J. A. Pollux, D. N. Reznick. 2011. Reproductive strategies in poeciliine fishes. In Evans, J., A. Pilastro and I. Schlupp (eds.). Ecology and Evolution of Poeciliid Fishes. The University of Chicago Press.

Pires, M. N., K.E. McBride, R. D. Bassar, J. U. Regus, T. Garland, Jr, and D. N. Reznick. 2011. Why do placentas evolve? An evaluation of the life-history facilitation hypothesis in the fish genus Poeciliopsis. Functional Ecology 25(4): 757-768.

Meredith, R. W., M. N. Pires, D. N. Reznick, M. S. Springer. 2011. Molecular phylogenetic relationships and the coevolution of placentotrophy and superfetation in Poecilia (Poeciliidae: Cyprinodontiformes). Molecular Phylogenetics and Evolution 59(1):148-57

Meredith, R. W., M. N. Pires, D. N. Reznick, M. S. Springer. 2010. A Molecular Phylogeny for Poecilia (Micropoecilia) (Poeciliidae: Cyprinodontiformes) Based on Nuclear and Mitochondrial DNA Sequences. Molecular Phylogenetics and Evolution 55: 631–639.

Pires, M. N., J. Arendt, D. N. Reznick. 2010. The evolution of placentas and superfetation in the fish genus Poecilia (Cyprinodontiformes: Poeciliidae: subgenera Micropoecilia and Acanthophacelus). Biological Journal of the Linnean Society. Biological Journal of the Linnean Society 99: 784–796

Pollux, B. J. A., M.N. Pires, A.I. Banet, D.N. Reznick. 2009. Evolution of Placentas in the Fish Family Poeciliidae: An Empirical Study of Macroevolution. Annual Review of Ecology, Evolution, and Systematics 40: 271-28

Turcotte, M. M., M. N. Pires, R. C. Vrijenhoek, and D. N. Reznick. 2008. Pre- and postfertilization maternal provisioning in livebearing fish species and their hybrids (Poeciliidae: Poeciliopsis). Functional Ecology 22:1118-1124.

Pires, M. N., K. E. McBride, and D. N. Reznick. 2007. Interpopulation variation in life-history traits of Poeciliopsis prolifica: implications for the study of placental evolution. Journal of Experimental Zoology Part A 307A:113-125. [Issue Cover Article; erratum published on May 2007]

Goodman, R. H., Jr., S. K. Watanabe, K. P. Condon, M. N. Pires, and M. S. Benton. 1998. Batrachoseps gabrieli (San Gabriel Mountain slender salamander). Herpetological Review 29:171.

Pires, M. N., and L. Seldin. 1997. Evaluation of Biolog system for identification of strains of Paenibacillus azotofixans. Antonie Van Leeuwenhoek International Journal of General and Molecular Microbiology 71:195-200.